Riccia

Riccia is a genus of liverworts in the order Marchantiales. These plants are small and thalloid, i.e. not differentiated into root, stem and leaf. Depending on species, the thallus may be strap-shaped and about 0.5 to 4 mm wide with dichotomous branches or may form rosettes or hemirosettes up to 3 cm in diameter, that may be gregarious and form intricate mats. The thallus is dorsiventrally differentiated. Its upper (dorsal) surface is green and chlorophyll-bearing, with a mid-dorsal longitudinal sulcus (furrow or groove). Air pores occasionally break through the dorsal surface, giving the thallus a dimpled appearance. In exceptional members such as Riccia caroliniana of Northern Australia and Riccia sahyadrica of Western Ghats, the photosynthetic region is confined to the lower half of the thallus. The lower (ventral) surface has a mid-ventral ridge bearing multicellular scales that originate as a single row but normally separate into two rows as the thallus widens. The scales are multicellular and hyaline (glassy) in appearance, or violet due to the pigment anthocyanin. Rhizoids are nearly lacking in aquatic forms, but there are usually numerous unicellular rhizoids of two types on the ventral surface.