Hydrachnidiae

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Hydrachnidia, also known as "water mites", Hydrachnidiae, Hydracarina or Hydrachnellae, are among the most abundant and diverse groups of benthic arthropods, composed of 6,000 described species from 57 families. As water mites of Africa, Asia, and South America have not been well-studied, the numbers are likely to be far greater. Other taxa of parasitengone mites include species with semi-aquatic habits, but only the Hydracarina are properly subaquatic. Water mites follow the general Parasitengona life cycle: active larva, inactive (calyptostasic) protonymph, active deutonymph, inactive tritonymph and active adult. Usually, larvae are parasites, while deutonymphs and adults are predators. Water mites may be brilliant red or orange in colour, unusual among freshwater invertebrates, but they also display more subtle blues, greens and yellows. They are also unusual among mites in some lineages having movable, internalized eye lenses sunk deep within the prosoma rather than being set on the surface of the cuticle. The group has two synapomorphies, features inherited from a common ancestor. In larvae, the genu of the palp has two setae. In post-larval stages, there are complex dermal structures consisting of a gland paired with a sensory seta (glandularia), possibly for defense against predators.