Prohercostomus

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Prohercostomus is an extinct genus of flies in the family Dolichopodidae, known from Baltic amber and Rovno amber from the Eocene. It was originally established as a subgenus of Hercostomus, but was later raised to genus rank. It includes 13 species described by Fernand Meunier between 1907 and 1908 that were originally placed in the genera Dolichopus, Gymnopternus and Poecilobothrus. In 1999, a new genus and species Sympycnites primaevus was described from a single female labelled as being from Early Cretaceous-aged Lebanese amber. This was considered unusual, because all other known fossils of Dolichopodidae in its strict sense (excluding the subfamilies Parathalassiinae and Microphorinae) were found in Cenozoic deposits. Dolichopodidae researchers later considered that the amber was probably mislabeled, and placed Sympycnites primaevus in synonymy with Prohercostomus noxialis in 2022.