Titanis

Titanis (meaning "Titan" for the Titans of Greek mythology) is a genus of phorusrhacid ("terror birds", a group originating in South America), an extinct family of large, predatory birds, in the order Cariamiformes that inhabited the United States from the early Pliocene to early Pleistocene. The first fossils were unearthed by amateur paleontologists/archaeologists Benjamin Waller and Robert Allen from the Santa Fe River in Florida and were named Titanis walleri by ornithologist Pierce Brodkorb in 1963, the species name honoring Waller. The holotype material is fragmentary, consisting of only an incomplete right tarsometatarsus (lower leg bone) and phalanx (toe bone), but comes from one of the largest phorusrhacid individuals known. In the years following the description, many more isolated elements have been unearthed from sites from other areas of Florida, Texas, and California. The species was classified in the subfamily Phorusrhacinae, which includes some of the last and largest phorusrhacids like Devincenzia and Kelenken. Like all phorusrhacids, Titanis had elongated hind limbs, a thin pelvis, proportionally small wings, and a large skull with a hooked beak. It was one of the largest phorusrhacids, possibly similar in size to Phorusrhacos based on preserved material. More recent estimates placed Titanis at 1.4 to 2 meters (4.6 to 6.6 ft) in height and over 300 kilograms (660 lb) in body mass. Due to the fragmentary fossils, the anatomy is poorly known, but several distinct characters on the tarsometatarsus have been observed. The skull is estimated to have been between 36 centimeters (14 in) and 56 centimeters (22 in) in length, one of the largest known from any bird. Phorusrhacids are thought to have been ground predators or scavengers, and have often been considered apex predators that dominated Cenozoic South America in the absence of placental mammalian predators, though they did co-exist with some...