Panperissodactyla

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Panperissodactyla ("all perissodactyls", alternatively spelled Pan-Perissodactyla) is a clade of ungulates containing living order Perissodactyla (odd-toed ungulates) and all extinct ungulates more closely related to Perissodactyla than to Artiodactyla (even-toed ungulates). Groups thought to belong to this clade include Anthracobunia (including the families Anthracobunidae and Cambaytheriidae) known from the Paleogene of the Indian subcontinent, as well as the South American native ungulate groups Litopterna and Notoungulata, both of which went extinct approximately 12,000 years ago. Other South American native ungulate groups also possibly belong to the clade, but their placement is uncertain. The enigmatic aquatic Desmostylia have also been suggested to be related to perissodactyls in some studies, though others recover them as members of afrotherian clade Tethytheria. The Northern Hemisphere "condylarth" group Phenacodontidae has been placed as closely related to perissodactyls in some studies, though others recover it as unrelated.